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Date of Award


Document Type


Degree Name

Master of Arts (M.A.)



First Advisor

Alden E. Noble


The genus Probolitrema was established by Looss (1901) in recognition of the location of vitellaria and multiple testes entirely outside the extra caecal field in Anaporrhutum richardii Lopez (1888) which differentiated it from A.albidum Ofenheim (1900) with the vitellaria within and the testes partly within and partly without the intra-caecal field. Ofenhein had created the genus Anaporrhutum for the above two species because of their multiple testes. A.albidum was a new species described by him; his A.richiardii Lopez specimens he considered to be the same species as Distomum richiardii Lopez, described in detail by Monticelli in 1893. Looss pointed out that they were not identical; Ofenheim’s specimens possessed succkers of nearly eual size while Monticelli’s specimens (which were probably D.richiardii Lopez) possessed a very large and powerful ventral sucker. Looss therefore retained P.richiardii Lopez as the type species of the genus and created a new species, P.capense, for A.richiardii Ofenheim.

ABout sixty individuals of Probolitrema tomalis were obtained from the coelom of a stingray, Dasyatis dipterura, on July 2, 1933, from Tomales Bay, California. No observations concerning the living animal were recorded.



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