Campus Access Only
All rights reserved. This publication is intended for use solely by faculty, students, and staff of University of the Pacific. No part of this publication may be reproduced, distributed, or transmitted in any form or by any means, now known or later developed, including but not limited to photocopying, recording, or other electronic or mechanical methods, without the prior written permission of the author or the publisher.
Date of Award
Thesis - Pacific Access Restricted
Master of Science (M.S.)
Eric O. Thomas
First Committee Member
Second Committee Member
C. Greg Anderson
Third Committee Member
Paul A. Richmond
Normal male dwarf African clawed frogs, Hymenochirus sp., possess bilateral, sexually dimorphic, subcutaneous breeding glands just posterior to the forelimbs. Previous studies have shown these glands release pheromones that attract conspecific females. This thesis shows the pheromones also stimulate the reproductive system of conspecific females. Females exposed to normal males prior to mating then allowed to mate with the normal males released a higher number of eggs than females who were not exposed to normal males prior to mating. Microscopic examination of ovarian tissue revealed that females exposed to normal males also produced more highly-developed oocytes than did females not exposed to normal males. These results suggest male Hymenochirus use pheromones not only to attract potential mates, but to increase female receptivity and readiness to mate. Evolutionarily, these pheromonal effects would likely benefit males by increasing their chances of breeding, increasing the number of eggs released by their mates, and thus the number of offspring in the next generation.
Madison, Amanda L.. (2005). Effects of male breeding gland in hymenochirus on female reproductive output. University of the Pacific, Thesis - Pacific Access Restricted. https://scholarlycommons.pacific.edu/uop_etds/610
To access this thesis/dissertation you must have a valid pacific.edu email address and log-in to Scholarly Commons.Find in PacificSearch
If you are the author and would like to grant permission to make your work openly accessible, please email